everything.study
BIOL 335

Limnology and Aquatic Ecology

3.00 units · Undergraduate

Fall 2025

Topics

  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish
  • Integrative Limnology and Applications (~13 lectures: Paleolimnology
  • Acidification
  • Shallow
  • Tropical lakes
  • Surprises in limnology
  • Lakes on Mars?)
  • Environmental change
  • Freshwater
  • Development
  • Learning
  • Ecosystems
  • Critical thinking
  • Environmental health

Readings

  • Listed material: The only material you will be responsible for is material covered in lectures, labs, and the field trip. However, detailed background material and many of the figures etc. used in lectures will be coming from the new textbook: Jones, I.D. and Smol, J.P. [Editors]. 2024. Wetzel’s Limnology: Lake and River Ecosystems. 4th Edition. Elsevier, Oxford. 1088 pp. https://shop.elsevier.com/books/wetzels-limnology/jones/978-0-12-822701-5
  • Listed material: Approximate Learning Hours
  • Listed material: Teaching method Average hours per week Number of weeks Total hours In-class hours Lecture ~3 12 ~36 Seminar
Fall 2024

Topics

  • Limnology
  • Ecology
  • Learning
  • Critical thinking
  • Ecosystems
  • Environmental health
  • Biodiversity

Readings

  • Listed material: 5 The textbook for this course Wetzel's Limnology: Lake and River Ecosystems (2023) - this text is available as an ebook from the Queen's Library (Wetzel's Limnology | ScienceDirect (queensu.ca)). This is an updated version of the classic limnology text by Wetzel (2021) - Limnology: Lake and River Ecosystems (3rd edition).
Fall 2023

Topics

  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~13 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • In-class hours
  • Group learning
  • Freshwater
  • Development
  • Ecosystems
  • Critical thinking
  • Environmental health

Readings

  • Listed material: There is no required textbook, although I will generally be following:
  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. The summaries at the end of each chapter are excellent.
  • Listed material: Bob Wetzel died shortly after completing the 3rd edition. Note this textbook will be replaced something this autumn by one I co-edited – in production at time of writing this syllabus:
  • Listed material: Jones, I. and Smol, J.P. [Editors]. 2024. (in production) Wetzel’s Limnology: Lake and River Ecosystems. 4th Edition. Elsevier, Oxford.
  • Listed material: Sunday Oct. 1).
  • Listed material: Approximate Learning Hours
  • Listed material: Teaching method Average hours per week Number of weeks Total hours In-class hours Lecture ~3 12 36 Seminar
  • Listed material: Laboratory 3 4 labs (see onQ page) and lab exam ~13 + field trip below Tutorial
  • Listed material: Group learning
  • Listed material: Individual instruction
  • Listed material: Other Online activity
  • Listed material: Off-campus activity Day trip to Queen’s University Biology Station
  • Listed material: Leave by 7:30am and return by ~6pm ~11 Private study
2022

Topics

  • Introduction to Limnology and Aquatic Ecology
  • Chemical Limnology (~7 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica
  • Case
  • Biological Limnology (~5 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~10+ lectures: Paleolimnology
  • Acidification
  • Lake remediation
  • Contaminants
  • Invasive species
  • Arctic and Antarctic lakes
  • Tropical lakes
  • Climate change
  • Multiples
  • In-class hours
  • Lecture/in class
  • Group learning
  • Ecosystems
  • Critical thinking
  • Environmental health

Readings

  • Listed material: Online versions are available for < Option A: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. The summaries at the end of each chapter are excellent. Option B: Kalff, J. 2022. Limnology: Inland Water Ecosystems, Prentice Hall. Several scientific journal articles relevant to the material presented in the course will be linked to the library via the BIOL335 OnQ website.
  • Listed material: Teaching method Average hours per week Number of weeks Total hours In-class hours Lecture/in class ~3 12 36 Seminar
  • Listed material: Laboratory 3 4 labs (see OnQ page) and lab exam ~13 + field trip (below) Tutorial
  • Listed material: Group learning
  • Listed material: Individual instruction
  • Listed material: Other Online activity
  • Listed material: Off-campus activity Day trip to Queen’s University Biology Station) Oct. 1 or 2 Leave 7:30 am, return by 6 pm. Private study
Fall 2021

Topics

  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~13 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. A number of scientific journal articles relevant to the material presented in the course will be posted on the BIOL335 website in PDF format. Most of these papers should also be available from the Queen’s library system.
2020-2021

Topics

  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~13 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. A number of scientific journal articles relevant to the material presented in the course will be posted on the BIOL335 website in PDF format. Most of these papers should also be available from the Queen’s library system.
2019-2020

Topics

  • methodology including chemical and biological sampling techniques, as well as plankton identification. In
  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~12 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. A number of scientific journal articles relevant to the material presented in the course will be posted on the BIOL335 website in PDF format. Most of these papers should also be available from the Queen’s library system.
2017-2018

Topics

  • methodology including chemical and biological sampling techniques, as well as plankton identification. In
  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~10 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. Reserve material: There is also a useful laboratory/field manual (Wetzel, R.G. and Likens, G.E. 1991. Limnological Analyses, Springer-Verlag, New York) on reserve in the library. Most of these papers should also be available from the Queen’s library system.
2016-2017

Topics

  • methodology including chemical and biological sampling techniques, as well as plankton identification. In
  • Introduction to Limnology and Aquatic Ecology
  • Chemical Limnology (~8 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Biological Limnology (~5 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~10-12 lectures: Paleolimnology
  • Acidification
  • Lake remediation
  • Contaminants
  • Invasive species
  • Arctic and Antarctic lakes
  • Tropical lakes
  • Climate change
  • Multiples
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. Reserve material: There is also a useful laboratory/field manual (Wetzel, R.G. and Likens, G.E. 1991. Limnological Analyses, Springer-Verlag, New York) on reserve in the library. Most of these papers should also be available from the Queen’s library system.
2015-2016

Topics

  • methodology including chemical and biological sampling techniques, as well as plankton identification. In
  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~10 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. Reserve material: There is also a useful laboratory/field manual (Wetzel, R.G. and Likens, G.E. 1991. Limnological Analyses, Springer-Verlag, New York) on reserve in the library. Most of these papers should also be available from the Queen’s library system.
2014-2015

Topics

  • methodology including chemical and biological sampling techniques, as well as plankton identification. In
  • Introduction to Limnology and Aquatic Ecology
  • Chemical Limnology (~8 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Biological Limnology (~5 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~10-12 lectures: Paleolimnology
  • Acidification
  • Lake remediation
  • Contaminants
  • Invasive species
  • Arctic and Antarctic lakes
  • Tropical lakes
  • Climate change
  • Multiples
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. Reserve material: There is also a useful laboratory/field manual (Wetzel, R.G. and Likens, G.E. 1991. Limnological Analyses, Springer-Verlag, New York) on reserve in the library. Most of these papers should also be available from the Queen’s library system.
2013-2014

Topics

  • methodology including chemical and biological sampling techniques, as well as plankton identification. In
  • Introduction to Limnology and Aquatic Ecology
  • Physical Limnology (~4 lectures: Lake morphometry
  • Light in lakes
  • Heat in lakes
  • Water movements)
  • Chemical Limnology (~ 6 lectures: Oxygen
  • Salinity
  • Carbon
  • Nitrogen
  • Phosphorus
  • Sulfur and Silica)
  • Origin of Lakes (1 lecture: Geographic aspects of limnology)
  • Biological Limnology (~6 lectures: Phytoplankton and primary production
  • Zooplankton
  • Fish and trophic
  • Integrative Limnology and Applications (~10 lectures: Paleolimnology
  • Acidification
  • Shallow lakes and
  • limnology
  • Lakes on Mars?)
  • Freshwater
  • Development

Readings

  • Listed material: Wetzel, R.G. 2001. Limnology: Lake and River Ecosystems (3rd edition), Academic Press. Reserve material: There is also a useful laboratory/field manual (Wetzel, R.G. and Likens, G.E. 1991. Limnological Analyses, Springer-Verlag, New York) on reserve in the library. Most of these papers should also be available from the Queen’s library system.
Sources